1105 RESEARCH REPORT INTRODUCTION Ovules are seed precursors and typically have two sheathing structures, termed the inner and outer integuments, that form the seed coat following fertilization. Numerous genes involved in integument patterning and growth have been identified from forward genetic screens (Skinner et al., 2004; Colombo et al., 2008). One such gene, ABERRANT TESTA SHAPE (ATS) encodes
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