1Laboratory of Neurophysiology, National Institute for Basic Biology, Okazaki, Japan 2Department of Basic Biology, The Graduate University for Advanced Studies (SOKENDAI), Miura, Japan 3Department of Psychology, Ritsumeikan University, Kyoto, Japan 4Department of Physiological Sciences, The Graduate University for Advanced Studies (SOKENDAI), Miura, Japan 5Division of Integrative Physiology, Natio