Density-dependent mating behaviors reduce male mating harassment in locusts View ORCID ProfileKoutaro Ould Maeno, View ORCID ProfileCyril Piou, Sidi Ould Ely, Sid’Ahmed Ould Mohamed, Mohamed El Hacen Jaavar, View ORCID ProfileSaïd Ghaout, and View ORCID ProfileMohamed Abdallahi Ould Babah Ebbe aLivestock and Environment Division, Japan International Research Center for Agricultural Sciences (JIRCA